cat

The cat (or domestic cat, house cat) is a small carnivorous mammal and a subspecies of the wild cat. The cat is a skilled predator and intelligent animal, known to hunt over 1,000 species for food, and capable of being trained (and learning by itself) to obey simple commands and manipulate simple mechanisms (see cat intelligence). It has lived in close association with humans for 3,500 to 9,500 years,^[1] figuring prominently in the mythology and legends of several cultures. In 2004, a grave was excavated on Cyprus that contained the skeletons, laid close to one another, of both a human and a cat. The grave is estimated to be 9,500 years old, pushing back the earliest known feline-human association significantly.[3]

Cats come in dozens of breeds and a variety of colors. Due to mutations, some are hairless and some are tailless. They use more than one hundred vocalizations and types of body language for communication, including calling ("meow" or "miaou"), purring, hissing, growling, chirping, clicking, and grunting.^[2] Like horses and other domesticated animals, cats can become feral, living effectively in the wild. Feral cats will often form small feral cat colonies. Animal welfare organizations note that few abandoned cats are able to survive long enough to become feral, most being killed by vehicles, or succumbing to starvation, predators, exposure, or disease.

Legends and myths about the cat exist in many cultures, from the ancient Egyptians and Chinese to the Vikings. They have been both revered and vilified by different cultures.

The trinomial name of the domestic cat is Felis silvestris catus; its closest pre-domesticated ancestor is believed to be the African wild cat, Felis silvestris lybica.^[3]

Scientific classification

The domestic cat was named Felis catus by Carolus Linnaeus in his Systema Naturae of 1758. Johann Christian Daniel von Schreber named the wild cat Felis silvestris in 1775. The domestic cat is now considered a subspecies of the wild cat: by the strict rule of priority of the International Code of Zoological Nomenclature the name for the species thus ought to be F. catus since Linnaeus published first. However, in practice almost all biologists use F. silvestris for the wild species, using F. catus only for the domesticated form.

In opinion 2027 (published in Volume 60, Part 1 of the Bulletin of Zoological Nomenclature, March 31 2003^[4]) the International Commission on Zoological Nomenclature "conserved the usage of 17 specific names based on wild species, which are predated by or contemporary with those based on domestic forms", thus confirming F. silvestris for the wild cat and F. silvestris catus for its domesticated subspecies. (F. catus is still valid if the domestic form is considered a separate species.)

Johann Christian Polycarp Erxleben named the domestic cat Felis domesticus in his Anfangsgründe der Naturlehre and Systema regni animalis of 1777. This name, and its variants Felis catus domesticus and Felis silvestris domesticus, are often seen, but they are not valid scientific names under the rules of the International Code of Zoological Nomenclature.

Nomenclature

A group of cats is referred to as a clowder, a male cat is called a tom, and a female is called a queen. The male progenitor of a cat, especially a pedigreed cat, is its sire, and its female progenitor is its dam. An immature cat is called a kitten (which is also an alternative name for young rats, rabbits, hedgehogs, beavers, squirrels and skunks). In medieval Britain, the word kitten was interchangeable with the word catling. A cat whose ancestry is formally registered is called a pedigreed cat, purebred cat, or a show cat (although not all show cats are pedigreed or purebred). In strict terms, a purebred cat is one whose ancestry contains only individuals of the same breed. A pedigreed cat is one whose ancestry is recorded, but may have ancestors of different breeds (almost exclusively new breeds; cat registries are very strict about which breeds can be mated together). Cats of mixed ancestry are referred to as domestic longhairs and domestic shorthairs or commonly as random-bred, moggies, mongrels, mutt-cats or alley cats. The ratio of pedigree/purebred cats to random-bred cats varies from country to country. However, generally speaking, purebreds are less than ten percent of the total feline population.^[5]

The word cat derives from Old English catt, which belongs to a group of related words in European languages, including Latin cattus, Byzantine Greek κάττα, Old Irish cat, and Old Church Slavonic kotka. The ultimate source of all these terms, however, is unknown.^[6]

The term puss (as in pussycat or Puss in boots) may come from Dutch (from "poes", a female cat, or the diminutive "poesje", an endearing term for any cat) or from other Germanic languages.

Characteristics

Physical features

Cats typically weigh between 2.5 and 7 kg (5.5–16 pounds); however, some breeds, such as the Maine Coon can exceed 11.3 kg (25 pounds). Some have been known to reach up to 23 kg (50 pounds) due to overfeeding. Conversely, very small cats (less than 1.8 kg - 4.0 lbs) [4] have been reported.

In captivity, indoor cats typically live 14 to 20 years, though the oldest-known cat lived to age 36.^[7] Domestic cats tend to live longer if they are not permitted to go outdoors (reducing the risk of injury from fights or accidents and exposure to diseases) and if they are spayed or neutered. Some such benefits are neutered male cats cannot develop testicular cancer, spayed female cats cannot develop ovarian cancer, and both have a reduced risk of mammary cancer.^[8] In contrast, feral cats living in modern urban environments often live only two years, or less. Feral cats in maintained colonies can live much longer; the British Cat Action Trust reported a 19-year-old feral female. The oldest feral cat was "Mark" who was maintained by the British charity Cats Protection and who reached 26 years of age.

Cats are athletic. Fast sprinters, they are capable of reaching speeds of about 50 km/h (about 30 mph) for short distances. They can often jump vertically up a fence or wall over 2 meters (over 7 feet) from stationary. The domestic cat is one of the few four-legged animals that do not possess rigid collarbones [5]. This allows a cat to fit through an opening about the size of its head.

Ears

Thirty-two individual muscles in the ear allow for a manner of directional hearing;^[9] the cat can move each ear independently of the other. Because of this mobility, a cat can move its body in one direction and point its ears in another direction. Most cats have straight ears pointing upward. Unlike dogs, flap-eared breeds are extremely rare. (Scottish Folds are one such exceptional genetic mutation.) When angry or frightened, a cat will lay its ears back, to accompany the growling or hissing sounds it makes. Cats will also turn their ears back when they are playing, or occasionally to show interest in a sound coming from behind them.

Metabolism

Cats conserve energy by sleeping more than most animals, especially as they grow older. Daily durations of sleep vary, usually 12–16 hours, with 13–14 being the average. Some cats can sleep as much as 20 hours in a 24-hour period. The term cat nap refers to the cat's ability to fall asleep (lightly) for a brief period and has entered the English lexicon – someone who nods off for a few minutes is said to be "taking a cat nap".

Due to their crepuscular nature, cats are often known to enter a period of increased hyperactivity and playfulness during the evening and early morning, dubbed the "evening crazies", "night crazies" or "mad half hour" by some.^[10][11]

The temperament of a cat can vary depending on the breed and socialization. Cats with "oriental" body types tend to be thinner and more active, while cats that have a "cobby" body type tend to be heavier and less active.

The normal body temperature of a cat is between 38 and 39 °C (101 and 102.2 °F).^[12] A cat is considered febrile (hyperthermic) if it has a temperature of 39.5 °C (103 °F) or greater, or hypothermic if less than 37.5 °C (100 °F). For comparison, humans have a normal temperature of approximately 36.8 °C (98.2 °F). A domestic cat's normal heart rate ranges from 140 to 220 beats per minute, and is largely dependent on how excited the cat is. For a cat at rest, the average heart rate should be between 150 and 180 bpm, about twice that of a human.

Legs

Cats, like dogs, are digitigrades: they walk directly on their toes, the bones of their feet making up the lower part of the visible leg. Cats are capable of walking very precisely, because like all felines they directly register; that is, they place each hind paw (almost) directly in the print of the corresponding forepaw, minimizing noise and visible tracks. This also provides sure footing for their hind paws when they navigate rough terrain.

Unlike dogs and most mammals, cats walk by moving both legs on one side and then both legs on the other. Most mammals move legs on alternate sides in sequence. Cats share this unusual gait with camels and a very few other mammals. There is no known connection between camels and cats which might explain this.

Like all members of family felidae, with the exception of the cheetah, cats have retractable claws. In their normal, relaxed position the claws are sheathed with the skin and fur around the toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hind feet. Cats can extend their claws voluntarily on one or more paws at will. Cats may extend their claws in hunting or self-defense, climbing, "kneading", or for extra traction on soft surfaces (bedspreads, thick rugs, etc.). The curved claws may become entangled in carpet or thick fabric, which may cause injury if the cat is unable to free itself.

Most cats have 4 claws at the rear and 5 at their front paws. But because of a rogue gene cats are prone to polydactyly, and some have 6 or 7 toes. The 5th front claw, the thumb, is on much higher position than those of the other fingers. Then even higher, there seems to be a 6th finger, but it is not. This special feature of the front paws, on the inside of the wrists, is the carpal pad, also found on the paws of big cats and dogs. It has no function in normal walking but is thought to be an anti-skidding device during jumping. If present, the 5th claw at the rear legs, corresponding with the big toe, is called the dew-claw.

Perching and falling

Most breeds of cat have a noted fondness for settling in high places, or perching. Animal behaviorists have raised a number of explanations, the most common being that height gives the cat a better observation point, allowing it to survey its "territory" and become aware of activities of people and other pets in the area. In the wild, a higher place may serve as concealed site from which to hunt.^[13] Height, therefore, can also give cats a sense of security and prestige.

This fondness for high spaces, however, can dangerously test the popular axiom that a cat "always lands on its feet." The American Society for the Prevention of Cruelty to Animals warns owners to safeguard the more dangerous perches in their homes, to avoid "high-rise syndrome," where an overconfident cat falls from an extreme height.^[14]

During a fall, a cat can reflexively twist its body and right itself using its acute sense of balance and flexibility.^[15] This is known as the cat's "righting reflex." It always rights itself in the same way, provided it has the time to do so during a fall. Thus, high (multi-story) falls can be less dangerous to them than those of only a few meters. Many cases are known of cats falling from substantial heights (5 to 10 stories) and surviving almost unscathed. Contrary to popular belief, cats without a tail also have this ability, since a cat mostly moves its hindlegs and relies on conservation of angular momentum to set up for landing, and the tail is in fact little used for this feat.^[16]

Senses

Measuring the senses of any animal can be difficult because there is usually no explicit communication (e.g., reading aloud the letters of a Snellen chart) between the subject and the tester.

While a cat's senses of smell may not be as keen as, say, that of a mouse, it is superior in many ways to those of humans. These along with the cat's highly advanced hearing, eyesight, taste, and touch receptors make the cat extremely sensitive among mammals.

Sight

Testing indicates that a cat's vision is superior at night in comparison to humans, and inferior in daylight. Cats, like dogs and many other animals, have a tapetum lucidum that reflects extra light to the retina. While this enhances the ability to see in low light, it appears to reduce net visual acuity, thus detracting when light is abundant. In very bright light, the slit-like iris closes very narrowly over the eye, reducing the amount of light on the sensitive retina, and improving depth of field. The tapetum and other mechanisms give the cat a minimum light detection threshold up to seven times lower than that of humans. Variation in color of cats' eyes in flash photographs is largely due to the interaction of the flash with the tapetum.

Average cats have a visual field of view estimated at 200°, versus 180° in humans, with a binocular field (overlap in the images from each eye) narrower than that of humans. As with most predators, their eyes face forward, affording depth perception at the expense of field of view. Field of view is largely dependent upon the placement of the eyes, but may also be related to the eye's construction. Instead of the fovea which gives humans sharp central vision, cats have a central band known as the visual streak. Cats can apparently differentiate among colors, especially at close range, but without appreciable subtlety.

Cats have a third eyelid, the nictitating membrane, which is a thin cover that closes from the side and appears when the cat's eyelid opens. This membrane partially closes if the cat is sick; although in a sleepy, content cat this membrane is often visible. If a cat chronically shows the third eyelid, it should be taken to a veterinary surgeon.

Cats have a wide variation in eye color, the most typical colors being golden, green and orange. Blue eyes are usually associated with the Siamese breed, but they are also found in white cats. If a white cat has two blue eyes, it is oftentimes deaf; however, orange eyes usually indicate the cat is free of hearing problems. White cats having one blue and one other-colored eye are called "odd-eyed" and may be deaf on the same side as the blue eye.^[17] This is the result of the yellow iris pigmentation rising to the surface of only one eye, as blue eyes are normal at birth before the adult pigmentation has had a chance to express itself in the eye(s).

Hearing

Humans and cats have a similar range of hearing on the low end of the scale, but cats can hear much higher-pitched sounds, up to 64 kHz, which is 1.6 octaves above the range of a human, and even 1 octave above the range of a dog.^[18] When listening for something, a cat's ears will swivel in that direction; a cat's ear flaps (pinnae) can independently point backwards as well as forwards and sideways to pinpoint the source of the sound. Cats can judge within three inches (7.5 cm) the location of a sound being made one yard (approximately one meter) away - this can be useful for localizing prey etc.

Smell

A domestic cat's sense of smell is about fourteen times as strong as a human's. Cats have twice as many smell-sensitive cells in their noses as people do, which means they can smell things we are not even aware of. Cats also have a scent organ in the roof of their mouths called the vomeronasal, or Jacobson's organ, which causes cats to become extremely sensitive towards open flames, fire, or burning objects. When a cat wrinkles its muzzle, lowers its chin, and lets its tongue hang a bit, it is opening the passage to the vomeronasal. This is called gaping, "sneering", or "flehming". Gaping is the equivalent of the Flehmen response in other animals, such as dogs, horses and big cats.

Touch

Cats generally have about a dozen whiskers in four rows on each upper lip^{citation needed]}, a few on each cheek, tufts over the eyes and bristles on the chin. Whiskers may also be found on the cat's inner "wrists", and there are similar hairs which make up the cat's eyebrows. The Sphynx (a nearly hairless breed) may have full length, short, or no whiskers at all.

Whiskers (technically called vibrissae) can aid with navigation and sensation. Whiskers may detect very small shifts in air currents, enabling a cat to know it is near obstructions without actually seeing them. The upper two rows of whiskers can move independently from the lower two rows for even more precise measuring.

It is thought that a cat may choose to rely on the whiskers in dim light where fully dilating the pupils would reduce its ability to focus on close objects. The whiskers also spread out roughly as wide as the cat's body making it able to judge if it can fit through an opening.

Whiskers are also an indication of the cat's attitude. Whiskers point forward when the cat is inquisitive and friendly, and lie flat on the face when the cat is being defensive or aggressive.

A cat has about twenty-four movable whiskers, twelve on each side of its nose (some cats may have more). Whiskers are more than twice as thick as ordinary hairs, and their roots are set three times deeper than hairs in a cat's tissue. Richly supplied with nerve endings, whiskers give cats extraordinarily detailed information about air movements, air pressure and anything they touch.

A scientific word for whiskers is vibrissae, a name that suggests their exquisite sensitivity to vibrations in air currents. As air swirls and eddies around objects, whiskers vibrate too. Cats use messages in these vibrations to sense the presence, size, and shape of obstacles without seeing or touching them. Whiskers are also good hunting tools. A cat whose whiskers have been damaged may bite the wrong part of a mouse it's attacking, indicating that signals from these delicate structures provide cats with vital information about the shape and activity of its prey - interestingly, whiskers also help cats detect scents.

Whiskers can also be a bother to a cat, especially when the cat tries to eat food out of a bowl. The end of the whiskers touching the side of the bowl transfer irritating sensations to its brain, making it hard for it to continue eating.

When a kitten is cleaned by its mother, she may chew off some or all of the whiskers.

Cats also have a number of reinforced hairs similar to whiskers on other parts of their bodies: over the eyes, on the chin and at the back of the legs. [6]

Taste

According to the December 8, 2005 issue of National Geographic, cats cannot taste sugary foods due to a faulty sweet receptor gene. Some scientists believe this is related to the cat's diet being naturally high in protein, though it is unclear whether it is a cause or result of it.

Communication

Further information: Cat communication

Hunting and diet

Relative to size, domestic cats are very effective predators. They ambush and dispatch vertebrate prey using tactics similar to those of leopards and tigers by pouncing; they then deliver a lethal neck bite with their long canine teeth that severs the victim's spinal cord, or asphyxiate it by crushing the windpipe. The domestic cat can hunt and eat about one thousand species—many big cats will eat fewer than 100. Although, theoretically, big cats can kill most of these species as well, they often do not due to the relatively low nutritional content that smaller animals provide for the effort. An exception is the leopard, which commonly hunts rabbits and many other smaller animals.

Even well-fed domestic cats will hunt and kill birds, mice, rats and other small animals in the vicinity. They often present such trophies to their owner. The motivation is not entirely clear, but friendly bonding behaviors are often associated with such an action. It is probable that cats in this situation expect to be praised for their symbolic contribution to the group. Some theories suggest that cats see their owners gone for long times of the day and assume they are out hunting, as they always have plenty of food available. It is thought that a cat presenting its owner with a dead animal thinks it's 'helping out' by bringing home the kill.^{citation needed]}

Due to their hunting behaviour, in many countries feral cats are considered pests. Domestic cats are occasionally also required to have contained cat runs or to be kept inside entirely, as they can be hazardous to locally endangered bird species. For instance, various municipalities in Australia have enacted such legislation. In some localities, owners fit their cat with a bell in order to warn prey of its approach. Sometimes, the bell has the unwanted effect of "training" the cat to be an even stealthier killer.^{citation needed]}

Cats have highly specialized teeth and a digestive tract suitable to the digestion of meat. The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently functions to shear meat like a pair of scissors. While this is present in canines, it is highly developed in felines. The cat's tongue has sharp spines, or papillae, useful for retaining and ripping flesh from a carcass. These papillae are small backward-facing hooks that contain keratin and assist in their grooming. Domesticated cats eat relatively little vegetable matter. It is quite common, however, for cats to occasionally supplement their carnivorous diets with small amounts of grass or other plant matter to help their digestive tract. Whereas bears and dogs commonly supplement their diet of meat with fruits, berries, roots, and honey when they can get them, cats prefer to feed mainly on meat. All felines, including the big cats, have a genetic anomaly that prevents them from tasting sweetness,^[19] which, more than likely, is related to their meat dominated eating habits, and almost certainly related to their aversion to fruits and berries. However, many domesticated cats are known to like vegetables.^[20] Also, the majority of brand-name cat foods are primarily grain based, often containing large amounts of corn or rice and supplemented with meat byproducts and minerals and vitamins. Cats are also known to munch on grass, leaves, shrubs and houseplants to regurgitate whatever is upsetting their stomach.

In captivity, cats cannot live on an unsupplemented vegetarian diet because they cannot synthesize several nutrients they need and that are absent or rare in plant food. This applies mainly to taurine, vitamin A (cats cannot convert the pro-vitamin A that is abundant in plants to vitamin A proper) and to certain fatty acids. The absence of taurine causes the cat's retina to slowly degenerate, causing eye problems and (eventually) irreversible blindness. This condition is called central retinal degeneration (CRD). Cow's milk is a poor source of taurine and adult cats are generally lactose intolerant. Lactose-free milk is perfectly safe, but still not a substitute for meat.

Some vegetarians, however, feed their cats a vegetarian diet, with a supplement containing these specific nutrients and others tailored to meet the needs of cats.^[21] Vegan pre-supplemented kibble is also available.^[22]

Some houseplants are harmful to cats. The leaves of the Easter Lily can cause permanent and life-threatening kidney damage to cats. Philodendron are also poisonous to cats. Cat Fancy has a full list of plants harmful to cats.

Some cats have a fondness for catnip. While they generally do not consume it, they will often roll in it, paw at it, and occasionally chew on it (as catnip is sensed by the cat's Vomeronasal organ). The effect is usually relatively short, lasting for only a few minutes. After two hours or less, susceptible cats gain interest again. Several other species of plants cause this effect, to a lesser degree.

Cats can be fussy eaters. This can happen when the vomeronasal, or Jacobson's, organ becomes accustomed to a specific food, or if the cats are spoiled by their owners, at which point the cat will reject any food that does not fit the pattern it is expecting. It is also known for cats to merely become bored with their given food and decide to stop eating until they are tempted into eating again. It is not unknown for them to starve themselves to death doing this; although it is extremely rare for a cat to deliberately starve itself to the point of injury, the sudden loss of weight can cause a fatal condition called hepatic lipidosis, a liver dysfunction which causes pathological loss of appetite. Additionally, cats have been known to develop a fondness for "people food" such as chicken, bread, French fries, pizza, ice cream, tomato soup, bacon, carrot juice, olives, mushrooms, and carnitas burritos, as well as cat diet exotica such as corn kernels and diced cantaloupe or cantaloupe skin. Many "people foods" are not good for cats; chocolate, for example, can be fatal due to the presence of theobromine (see theobromine poisoning), although few cats will eat chocolate. Paracetamol, found in many familiar over-the-counter pharmaceutical products such as Tylenol, is extremely toxic to cats. Because they naturally lack enzymes needed to digest it, even minute portions of doses safe for humans can be fatal.^[23] Any suspected ingestion warrants immediate veterinary attention.^[24]

Also, a diet consisting of people food or unlimited access to normal cat food often leads to the cat becoming overfed and overweight. This is very unhealthy for the cat, leading to other health complications, such as diabetes, especially in neutered males. Such health conditions can be prevented through diet and exercise (playing), especially for cats living exclusively indoors.

Cats can also develop pica. Pica is a condition in which animals chew or eat unusual things such as fabric, plastic or wool. In cats, this is mostly harmless as they do not digest most of it, but can be fatal or require surgical removal if a large amount of foreign material is ingested (for example, an entire sock). It tends to occur more often in Siamese, Burmese, and breeds with these in their ancestry.

Domestic cats, especially young kittens, are known for their love of string play. Many cats cannot resist a dangling piece of string, or a piece of rope drawn randomly and enticingly across the floor. This notorious love of string is often depicted in cartoons and photographs, which show kittens or cats playing with balls of yarn. It is probably related to hunting instincts, including the common practice of kittens practice hunting of their mother's, and each other's, tail. If string is ingested, however, it can become caught in the cat’s stomach or intestines, causing illness, or in extreme cases, death. Due to the possible complications of ingesting string, string play is sometimes replaced with a laser pointer's dot, which some cats will chase. Some also discourage the use of laser pointers for pet play, however, because of the potential damage to sensitive eyes and/or the possible loss of satisfaction associated with the successful capture of an actual prey object, play or real. ^{citation needed]}

Because of their small size, domestic cats pose almost no danger to humans — the main hazard is the possibility of infection (e.g., cat scratch disease, or, rarely, rabies) from a cat bite or scratch. Cats can also potentially inflict severe scratches or puncture an eye, though this is quite rare. Dogs have been known to be blinded by cats in fights, in which the cat specifically targeted the eyes of the larger animal with some accuracy.

Cats can be destructive to ecosystems in which they are not native and whose species have not had time to adapt to their introduction. In some cases, cats have contributed to or caused extinctions -— for example, see the case of the Stephens Island Wren.

Hygiene

Cats are known for their fastidious cleanliness. They groom themselves by licking their fur, employing their hooked pappilae and saliva. Their saliva is a powerful cleaning agent, but it can provoke allergic reactions in humans. Some people who are allergic to cats—typically manifested by hay fever, asthma or a skin rash —quickly acclimate themselves to a particular animal and live comfortably in the same house with it, while retaining an allergy to cats in general.^{citation needed]} Many cats also enjoy grooming humans or other cats. Some cats occasionally regurgitate hair balls of fur that have collected in their stomachs as a result of their grooming. Longhair cats are more prone to this than shorthairs. Hairballs can be prevented with certain cat foods and remedies that ease elimination of the hair and regular grooming of the coat with a comb or stiff brush. Cats expend nearly as much fluid grooming as they do urinating.

Indoor cats may be provided a litter box containing sand or similar material (litter). This arrangement serves the same purpose as a toilet for humans. It should be cleaned daily and changed often (depending on the number of cats in a household and the type of litter; clumping litter stays cleaner longer, but has been reported to cause health problems in some cats.^[25]) A litterbox is recommended for indoor-outdoor cats as well. Litterboxes may pose a risk of toxoplasmosis transmission to susceptible pregnant women and immuno-compromised individuals. Transmission risk may be reduced by daily litterbox cleaning by someone other than child-bearing women.

In addition, some cats can be toilet trained, eliminating the litterbox and its attendant expense and smell. Training involves two or three weeks of incremental moves, such as moving and elevating the litterbox until it is near the toilet. For a short time, an adapter, such as a bowl or small box, may be used to suspend the litter above the toilet bowl. When training is complete, the cat uses the toilet by perching over the bowl.^[26]

Indoor cats will also benefit from being provided with a scratching post so they are less likely to ruin furniture with their claws.^[27] Some indoor cats, especially those that were taken as kittens from feral colonies, may not understand the concept of a scratching post, and as such they will ignore it. Nails can be trimmed, as with a small pair of electrical dikes (ie, cutting pliers), but care should always be taken to avoid cutting a vein in the quick of the claw.

Declawing

Main article: Onychectomy

Declawing is a major surgery known as onychectomy, performed under anesthesia, that removes the tip of each digit (from the first knuckle out) of the cat's forepaws (and sometimes the hind paws). There is a slight chance of death in the surgery, and a declawed cat may have an increased risk of infection and life-long discomfort in its paws. This surgery is not recommended for an adult animal and is considered an act of animal cruelty in many Western countries (see below).

People generally have cats declawed to prevent them from hunting and from damaging furniture. Rarely, vicious cats are declawed. In the United States, some landlords require that tenants' cats be declawed.

Veterinarians are generally critical of the procedure and some refuse to perform it because the absence of claws in a cat:

1. Deprives it of its main defense abilities, including escaping from predators by climbing trees;
2. Impairs its stretching and exercise habits, leading to muscle atrophy;
3. Compromises its ability to balance on thin surfaces such as railings and fence tops, leading to injury from falls;
4. Can cause insecurity and a subsequent tendency to bite.

This operation is rare outside of North America. In Finland, Germany, the Netherlands and Switzerland, declawing is forbidden by the laws against cruelty to animals.^[28] In many other European countries, it is forbidden under the terms of the European Convention for the Protection of Pet Animals, unless "a veterinarian considers [such] non-curative procedures necessary either for veterinary medical reasons or for the benefit of (the) animal".^[29] In Britain, animal shelters find it difficult to place imported cats that have been declawed and subsequently most are euthanized. In 2003, West Hollywood, California became the first U.S. jurisdiction to outlaw declawing by veterinarians or animal groomers practicing in city limits.

An alternative to declawing is the application of blunt, vinyl nail caps that are affixed to the claws with nontoxic glue, requiring periodic replacement when the cat sheds its claw sheaths (about every four to six weeks). However, the cat will still experience difficulties because the capped nails are not as effective as claws.

After a cat has been declawed, it should be allowed to rest, and restrained from jumping (if possible) for a few days. After being neutered/spayed/declawed (as with after any surgery), the cat will go through a period when being played with or picking it up may hurt it. This usually lasts a week or so, sometimes less. While some people suggest cats not be declawed until 5-6 months of age, many veterinarians who practice this surgery are of the opinion the cat can be declawed as soon as they are old enough to sustain surgery (around 2-3 months of age, depending on size). The reasoning is that younger cats are more adaptable to the amputation, and that distal phalanges in the cat at this age are still flexible cartilage rather than bone, may make the operation somewhat less painful.

Environment

The wild cat, ancestor of the domestic cat, is believed to have evolved in a desert climate, as evident in the behavior common to both the domestic and wild forms. Wild cats are native to all continents other than Australasia and Antarctica. Their feces are usually dry, and cats prefer to bury them in sandy places. Urine is highly concentrated, which allows the cat to retain as much fluid as possible. They are able to remain motionless for long periods, especially when observing prey and preparing to pounce. In North Africa there are still small wildcats that are probably related closely to the ancestors of today's domesticated breeds.

Cats enjoy heat and solar exposure, often sleeping in a sunny area during the heat of the day. Cats prefer warmer temperatures than humans do. People start to feel uncomfortable when their skin's temperature gets higher than about 44.5 °C (112 °F), but cats don't start to show signs of discomfort until their skin reaches about 52 °C (126 °F).

Being closely related to desert animals, cats can easily withstand the heat and cold of a temperate climate, but not for extended periods. Although certain breeds such as the Norwegian Forest Cat and Maine Coon have developed heavier coats of fur than other cats, they have little resistance against moist cold (eg, fog, rain and snow) and struggle to maintain their 39 °C (102 °F) body temperature when wet.

Most cats dislike immersion in water; one major exception is the Turkish Van breed which has an unusual fondness for water [7]. Abyssinians are also reported to be more tolerant of water than most cats.

Reproduction and genetics

Cats are seasonally polyestrous, which means they may have many periods of heat over the course of a year. A heat period lasts about 4 to 7 days if the female is bred; if she is not, the heat period lasts longer.

The male cat's penis has spines which point backwards. Upon withdrawal of the penis, the spines rake the walls of the female's vagina. The female needs this stimulation for ovulation to begin. Because this does not always occur, females are rarely impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, meaning different kittens in a litter may have different fathers.

The reproduction process can be very loud, as both cats vocalize loudly. If one is not used to the sounds of cats mating, it sounds very much like a cat fight.

The gestation period for cats is approximately 63-65 days. The size of a litter averages three to five kittens, with the first litter usually smaller than subsequent litters. Kittens are weaned at between six and seven weeks, and cats normally reach sexual maturity at 4-10 months (females) and to 5-7 months (males).

Cats are ready to go to new homes at about 12 weeks old (the recommended minimum age by Fédération Internationale Féline), or when they are ready to leave their mother. Cats can be surgically sterilized (spayed or neutered) as early as 6-8 weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as territory marking (spraying urine) in males and yowling (calling) in females. If an animal is neutered after such behavior has been learned, however, it may persist.

There are several genes responsible for the hair color identified. The combination of them gives different phenotypes. See Cat coat genetics.

Features like hair length, lack of tail or presence of a very short tail (bobtail cat) are also determined by single alleles and modified by polygenes.

The Cat Genome Project, sponsored by the Laboratory of Genomic Diversity at the U.S. National Cancer Institute Frederick Cancer Research and Development Center in Frederick, Maryland, focuses on the development of the cat as an animal model for human hereditary disease, infectious disease, genome evolution, comparative research initiatives within the family Felidae, and forensic potential.

Domestication

Like some other domesticated animals, cats live in a mutualistic arrangement with humans. It is believed that the benefit of removing rats and mice from humans' food stores outweighed the trouble of extending the protection of a human settlement to a formerly wild animal, almost certainly for humans who had adopted a farming economy. Unlike the dog, which also hunt and kill rodents, the cat did not eat grains, fruits, or vegetables. A cat that is good at hunting rodents is referred to as a mouser.

The venerable simile "like herding cats" refers to the seeming intractability of the ordinary house cat to training in anything, unlike dogs. Despite cohabitation in colonies, cats are lone hunters. It is no coincidence that cats are also "clean" animals; the chemistry of their saliva, expended in frequent grooming, appears to be a natural deodorant. If so, the function of this cleanliness is to decrease the chance a prey animal will notice the cat's presence in time. In contrast, dog's odour is an advantage in hunting, for a dog is a pack hunter; part of the pack stations itself upwind, and its odour drives prey towards the rest of the pack stationed downwind. This requires a cooperative effort, which in turn requires communications skills. No such communications skills are required of a lone hunter. It is likely this is part of the reason interacting with such an animal is problematic; cats in particular are labeled as opaque or inscrutable, if not obtuse, as well as aloof and self-sufficient. However, cats can be very affectionate towards their human companions, especially if they imprint on them at a very young age and are treated with consistent affection.

Human attitudes toward cats vary widely. Some people keep cats for companionship as pets. Others go to great lengths to pamper their cats, sometimes treating them almost as if they were children. When a cat bonds with its owner, the cat may, at times, display behaviors similar to that of a human. Such behavior may include a trip to the litter box before bedtime or snuggling up close to its companion in bed or on the sofa. Other such behaviors could include mimicking sounds of the owner or using certain sounds the cat picks up from the human; sounds representing specific needs of the cat, which the owner would recognize, such as a specific tone of meow along with eye contact that may represent "I'm hungry." The cat may also be capable of learning to communicate with the human using non-spoken language or body language such as rubbing for affection (confirmation), facial expressions and making eye contact with the owner if something needs to be addressed (e.g., finding a bug crawling on the floor for the owner to get rid of). Some owners like to train their cat to perform "tricks" commonly exhibited by dogs such as jumping; this is rare, however.

Allergies to cat dander are one of the most common reasons people cite for disliking cats. However, in some instances, humans find the rewards of cat companionship outweigh the discomfort and problems associated with these allergies. Many choose to cope with cat allergies by taking prescription allergy medicine and bathing their cats frequently, since weekly bathing will eliminate about 90% of the cat dander present in the environment. Recent studies have indicated that humans who are exposed to cats or dogs within the first year of their lives develop few animal allergies, while most adults who are allergic to animals did not have a cat or a dog as a pet in childhood ^{citation needed]}.

In urban areas, some people find feral and free-roaming pet cats annoying and intrusive. Unaltered animals can engage in persistent nighttime calling (termed caterwauling) and defecation or "marking" of private property. Indoor confinement of pets and TNR (trap, neuter, return) programs for feral cats can help; some people also use cat deterrents to discourage cats from entering their property.

In rural areas, farms often have dozens of semi-feral cats. Hunting in the barns and the fields, they kill and eat rodents that would otherwise spoil large parts of the grain crop. Many pet cats successfully hunt and kill rabbits, rodents, birds, lizards, frogs, fish, and large insects by instinct, but might not eat their prey. They may even present their kills, dead or maimed, to their humans, perhaps expecting them to praise or reward them, or possibly even to complete the kill and eat the mouse. Others speculate that the behavior is a part of the odd relationship between human and cat in which the cat is sometimes a 'kitten' (playing, being picked up and carried) and sometimes an adult (teaching these very large and peculiar kittens how to hunt by demonstrating what the point of it all is).

Despite its reputation as a solitary animal, the domestic cat is social enough to form colonies, but does not attack in groups as do lions. Some breeds like bengal, ocicat and manx are very social, but these breeds are exceptions. While each cat holds a distinct territory (sexually active males having the largest territories, and neutered cats having the smallest), there are "neutral" areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually aggressively chase away stranger cats, at first by staring, hissing, and growling, and if that does not work, by short but noisy and violent attacks. Fighting cats make themselves appear more impressive and threatening by raising their fur and arching their backs, thus increasing their visual size. Cats also exhibit this behavior while playing. Attacks usually comprise powerful slaps to the face and body with the forepaws as well as bites, but serious damage is rare; usually the loser runs away with little more than a few scratches to the face, perhaps the ears. Normally, serious negative effects will be limited to possible infections of the scratches and bites; though these have been known to sometimes kill cats if untreated. Sexually active males will usually be in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to the ears and nose. Not only males will fight; females will also fight over territory or to defend their kittens, and even neutered cats will defend their (smaller) territories aggressively.

Feral cats

Main article: Feral cat

Feral cats may live alone, but most are found in large groups called feral colonies with communal nurseries, depending on resource availability. Some lost or abandoned pet cats succeed in joining these colonies, probably for lack of an alternative. The average lifespan of such feral cats is much shorter than a domestic housecat, which can live sixteen years or more. Urban areas in the developed world are not friendly, nor adapted, environments for cats; most domestic cats are descended from cats in desert climates and were distributed throughout the world by humans. Nevertheless, some feral cat colonies are found in large cities, e.g., around the Colosseum and Forum Romanum in Rome.

Although cats are adaptable, feral felines are unable to thrive in extreme cold and heat, and with a very high protein requirement, few find adequate nutrition on their own in cities. In addition, they have little protection or understanding of the dangers from dogs, coyotes, and even automobiles. However, there are thousands of volunteers and organizations that trap these unadoptable feral felines, spay or neuter them, immunize the cats against rabies and feline leukemia, and treat them with long-lasting flea products. Before release back into their feral colonies, the attending veterinarian often nips the tip off one ear to mark the feral as spayed/neutered and inoculated, as these cats will more than likely find themselves trapped again. Volunteers continue to feed and give care to these cats throughout their lives, and not only is their lifespan greatly increased, but behavior and nuisance problems, due to competition for food, are also greatly reduced. In time, if an entire colony is successfully spayed and neutered, no additional kittens are born and the feral colony disappears. Many hope to see an end to urban feral cat colonies through these efforts.

Environmental issues

There are two divergent views about cats’ relationship with the natural environment.

• The first says: The environmental impact of feral cat programs and of indoor/outdoor cats is a subject of debate. Part of this stems from humane concern for the cats themselves and part arises from concerns about cat predation on endangered species. Nearly all studies agree that abandoned animals lead hard lives. Owners who can no longer keep their cats would do best to give them to friends, rescue organizations, or shelters. The amount of ecological damage done by indoor/outdoor cats depends on local conditions. The most severe impact occurs with island ecologies. Serious concerns also exist in places such as Florida where housecats are not native, where several small-sized endangered species live near human populations, and where the climate allows cats to breed throughout the year. Environmental concerns may be minimal in most of the UK where cats are an established species and few to none of the local prey species are endangered. Pet owners can contact veterinarians, ecological organizations, and universities for opinions about whether local conditions are suitable for outdoor cats. Additional concerns include potential dangers from larger predators and infectious diseases. Coyotes kill large numbers of housecats in the Southwestern United States, even in urban zones. FELV (feline leukemia), FIV (feline immunodeficiency virus), or rabies may be present in the area. If faced with conflicting evidence, the safe choice is to keep a cat indoors. Experts recommend a gradual transition to indoor life for cats who are accustomed to going outside.
• Those opposing this view stress that this allegation has never been proved. They say that damaging effects do not follow automatically from the fact that cats are predators. They point out that cats have played a useful role in vermin control for centuries, and that for many animals, especially in urban areas, cats are the only animal available to fill the vital role of predator. Without cats these species would overpopulate.

Overpopulation

According to the Humane Society of the United States, 3-4 million cats and dogs are euthanized each year in the United States and many more are confined to cages in shelters because there are significantly more animals being born than there are homes. Spaying or neutering pets helps keep the overpopulation down. [8] Local humane societies, SPCA's and other animal protection organizations urge people to spay or neuter their pets and to adopt animals from shelters instead of purchasing them.

Varieties of domestic cat

The list of cat breeds is quite large: most cat registries recognize between 35 and 70 breeds of cats, and several more are in development, with one or more of new breeds being recognized each year on average, having distinct features and heritage. The owners and breeders of show cats compete to see whose animal bears the closest resemblance to the "ideal" definition of the breed (see selective breeding). Due to common crossbreeding in populated areas, many cats are simply identified as belonging to the homogeneous breeds of domestic longhair and domestic shorthair, depending on their type of fur. In the United Kingdom and Australia, non-purebred cats are referred in slang as moggies (derived from "Maggie", short for Margaret, reputed to have been a common name for cows and calves in 18th-century England and latter applied to housecats during the Victorian era).^[30] In the United States, a non-purebred cat is sometimes referred to in slang as a barn or alley cat, even if it is not a stray.

Cats come in a variety of colors and patterns. These are physical properties and should not be confused with a breed of cat.

Household cats are divided into:

• Domestic longhaired
• Domestic shorthaired

Coat patterns

Cat coat genetics can produce a variety of coat patterns. Some of the most common are:

Bicolor, Tuxedo and Van 

This pattern varies between the tuxedo cat which is mostly black with a white chest, and possibly markings on the face and paws/legs, all the way to the Van pattern (so named after the Lake Van area in Turkey, which gave rise to the Turkish Van breed), where the only colored parts of the cat are the tail (usually including the base of the tail proper), and the top of the head (often including the ears). There are several other terms for amounts of white between these two extremes, such as harlequin or jellicle cat. Bicolor cats can have as their primary (non-white) color black, red, any dilution thereof and also tortoiseshell (see below for definition).

Tabby cat 

Striped, with a variety of patterns. The classic "blotched" tabby (or "marbled") pattern is the most common and consists of butterflies and bullseyes. The "mackerel" or "striped" tabby is a series of vertical stripes down the cat's side (resembling the fish). This pattern broken into spots is referred to as a "spotted" tabby. Finally, the tabby markings may look like a series of ticks on the fur, thus the "ticked" tabby, which is almost exclusively associated with the Abyssinian breed of cats. The worldwide evolution of the cat means that certain types of tabby are associated with certain countries; for instance, blotched tabbies are quite rare outside NW Europe, where they are the most common type.

Tortoiseshell and Calico

This cat is also known as a Calimanco cat or Clouded Tiger cat, and by the nickname "tortie". In the cat fancy, a tortoiseshell cat is randomly patched over with red (or its dilute form, cream) and black (or its dilute blue) mottled throughout the coat. Additionally, the cat may have white spots in its fur, which will make it a "tortoiseshell and white" cat or, if there is a significant amount of white in the fur and the red and black colors form a patchwork rather than a mottled aspect, the cat will be called a "calico". All calicos are tortoiseshell (as they carry both black and red), but not all tortoiseshells are calicos (which requires a significant amount of white in the fur and patching rather than mottling of the colors). The calico is also sometimes called a "tricolor cat". The Japanese refer to this pattern as mi-ke (meaning "triple fur"), while the Dutch call these cats a lapjeskat (meaning "patches cat"). A true tricolor must consist of three colors: a reddish color, dark or light; white; and one other color, typically a brown, black or blue, as described by American breeder Barbara French, writing for the Cat Fanciers community.^[31] Both tortoiseshell and calico cats are typically female because the coat pattern is the result of differential X chromosome inactivation in females (which, as with all normal female mammals, have two X chromosomes). Those male tortoiseshells that are created are usually sterile; conversely, cats where the overall color is ginger (orange) are commonly male (roughly in a 3:1 ratio). In a litter sired by a ginger tom, the females will be tortoiseshell or ginger. See "Tortoiseshell and Tricolour Cats" for an extensive genetic explanation for tricolor cats, and detailing the possible combinations of coloring.^[32]

Colorpoint

A flame point Siamese mix.

The colorpoint pattern is most commonly associated with Siamese cats, but also may appear in any domestic cat. A colorpoint cat will have dark colors on the face, ears, feet, and tail, with a lighter version of the same color on the rest of the body and possibly some white. The exact name of the colorpoint pattern depends on the actual color, so there are seal points (dark brown), chocolate points (warm lighter brown), blue points (dark gray), lilac points (silvery gray-pink), flame points (orange), and tortie (tortoiseshell mottling) points, among others. It should also be noted that colorpoint cats tend to darken with age, and the fur over a significant injury may sometimes darken or lighten depending on circumstances due to the fact that pigment synthesis in the fur is temperature-sensitive.

Body types

Cats can also come in several body types, ranging between two extremes:

Oriental 

Not a specific breed, but any cat with an elongated slender build, almond-shaped eyes, long nose, large ears (the Siamese and oriental shorthair breeds are examples of this).

Cobby 

Any cat with a short, muscular and compact build, roundish eyes, short nose, and small ears. The Persian breed is a prime example of a cobby cat.

History and mythology

Main article: History of cats

Cats have been kept by humans since at least ancient Egypt. In ancient Egypt, the cat god, Bast, was a goddess of the home and of the domestic cat, although she sometimes took on the war-like aspect of a lioness. She was the daughter of the sun god Ra, although she was sometimes regarded as the daughter of Amun. She was the wife of Ptah and mother of the lion-god Mihos. Her cult was centered on her sanctuary at Bubastis in the Nile Delta, where a necropolis has been found containing mummified cats. Bast was also associated with the "eye of Ra," acting as the instrument of the sun god's vengeance. She was depicted as a cat or in human form with the head of a cat, often holding the sacred rattle known as the sistrum.

It has been speculated that because of circumstantial evidence (to date unproved by DNA testing) cats resident on Kenya's Islands in the Lamu Archipelago may be the last living direct descendants of the sacred cats of ancient Egypt. The suggestive evidence is: similar body conformation, archeologically proven history of 1000 years habitation, ancient Red Sea trade between Lamu and Egypt, genetic insular isolation. ^{citation needed]}

According to Norse legend, Gleipnir (the fine ribbon used to bind Fenrir) was crafted by dwarfs from, among other items, the sound a cat makes when walking.

Several ancient religions believe that cats are exalted souls, companions or guides for humans, that they are all-knowing but are mute so they cannot influence decisions made by humans.

Muezza (Arabic: ﻣﻮﻴﺰا‎ ​) was the Prophet Muhammad's favorite cat. The most famous story about Muezza recounts how the call to prayer was given, and as Muhammad went to put on one of his robes, he found his cat sleeping on one of the sleeves, and instead of disturbing the cat he cut off the sleeve and let him sleep. When he returned, Muezza awoke and bowed down to Muhammad and in return he stroked him three times.

It is common lore that cats have nine lives. It is a tribute to their perceived durability, their occasional apparent lack of instinct for self-preservation, and their seeming ability to survive falls that would be fatal to other animals.

A medieval King of Wales, Hywel Dda (the Good) passed legislation making it illegal to kill or harm a cat. Other cultures of the time considered them evil, unlucky, or the consorts of witches.

References

See also

• Fred the Undercover Kitty
• Creme Puff
• Morris the Cat
• Simon (cat)
• Socks (cat)
• Spot (Star Trek)
• Hypoallergenic cat
• Felidae
• Big cat
• Cat body language
• Catnip
• Cat flap
• Cat Fanciers' Association
• Cats in ancient Egypt
• Kitten
• List of historical cats
• List of fictional cats
• Polydactyl cat (extra toes)
• Cat types
• Cat breeds

External links

Medical issues

• Feline Medical & Behaviour Database (large number of short articles)
• Ailurophilia Gone Bad
• "DNA Offers New Insight Concerning Cat Evolution", The New York Times, January 6, 2006.
• High-Resolution Images of the Cat Brain
• Onions are Toxic to Cats
• Information about the third eyelid of cats, and the problem of Kertao, or "dry eye".

Miscellaneous

• New Potential Smallest Cat
• BBC.CO.UK Description of the Cat

• Cationary: Meaningful Portraits of Cats by Sharon Montrose, ISBN 0-670-03059-7